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Tier I Right click to recruit a familiar to an empty cell for 5 coins.
Tier II Whenever you collect loot, upgrade all nearby familiars to strong familiars.
Tier III [2 Charges] Pass 10 turns.
Unlock I Unlock by having familiars
Unlock II Unlock II by having familiars
Unlock III Unlock III by having familiars

The Commander Mastery is focused on using Familiars.