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Tier I Right click to recruit a familiar to an empty cell for 5 coins.
Tier II Whenever you collect loot, upgrade all nearby familiars to strong familiars.
Tier III [2 Charges] Pass 10 turns.
Unlock I Unlock by having 10 or more familiars
Unlock II Unlock II by having 15 or more familiars
Unlock III Unlock III by having 30 or more familiars

The Commander Mastery is focused on using Familiars.