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Tier I Each stage has a burning cell. Burning cells do not hurt you.
Tier II Stages have extra burning cells equal to the number of stage mods.
Tier III [4 Charges] Shoot flares in all directions from visible burning cells.
Unlock I Unlock by having 10 simultaneous flares.
Unlock II Unlock II by having 20 simultaneous flares.
Unlock III Unlock III by having 30 simultaneous flares.

The Firefly Mastery is focused on burning cells.